In an alternative 1919, two special cats named Jet and Alley join together by chance and travel to "Americana", to live a better life at Pittsburgh. However, they're thrown into a case involving a missing cat named Jasmine. This mystery makes them embark on a perilous journey to find her. Faced with adversaries, they discover a sinister plot behind her abduction. In addition, their new and unexpected bond eventually forms an uncharted path amidst an advanced society fueled by science, corruption, and power.