1876. Among the filth and depravity of Covent Garden's flower markets, orphaned Irish sisters Flora and Rosie Flynn sell posies of violets and watercress to survive. It is a pitiful existence, made bearable only by each other's presence. When they become separated, the decision of a desperate woman sets their lives on very different paths. 1912. Twenty-one-year-old Tilly Harper leaves the peace and beauty of her native Lake District for London to become assistant housemother at one of Mr. Shaw's Training Homes for Watercress and Flower Girls. For years, the homes have cared for London's orphaned and crippled flower girls, getting them off the streets. For Tilly, the appointment is a fresh start, a chance to leave her troubled past behind. Soon after she arrives at the home, Tilly finds a notebook belonging to Flora Flynn. Hidden between the pages she finds dried flowers and a heartbreaking tale of loss and separation as Flora's entries reveal how she never stopped looking for her lost sister. Tilly sets out to discover what happened to Rosie—but the search will not be easy. Full of twists and surprises, it leads the caring and determined young woman into unexpected places, including the depths of her own heart.