A talented graffiti artist has been vandalizing the Church of Peace, but there have been no leads. Then a bomb destroys half the church, and its pastor is found dead beneath the altar. The evidence is misleading, and Det. Jesse Quinn's tunnel-visioned partner is eager to arrest Jesse's next-door neighbor. With her regular partner on family leave, Jesse is temporarily paired with her nemesis, a detective of the St John's County Sheriff's Office who thinks women don't belong on the Major Crimes squad. Zack Oliver is in a hurry to override her objections, close the case and move on. Jesse just needs to find the real killer of the popular local pastor before Zack pins the murder on an innocent man. Meanwhile, at home, Jesse is trying to reconcile with her teenage daughter who is grounded for entertaining a boy in her bedroom. The teen is now lobbying to move into Jesse's ex's house where she thinks the rules will be more relaxed. And then, there's Seth Cameron who would like to become more in Jesse's life than just a friend. When did her life get so complicated?