The story is set in Burgdale, a small Germanic settlement in a valley at the foot of a mountain range, and the neighbouring woodlands, pastures and dales. The area is inhabited by the interdependent Dalemen, who are weavers, smiths, and traders, the Woodlanders, who are hunters and carpenters, and the Shepherds. Their society is challenged by disruptions from the outside world in the form of the Sons of the Wolf, the descendants of the Wolfings from the previous novel, and the invading Dusky Men (the Huns). The Sons of the Wolf, driven from their original country by the Dusky Men, continue to resist the invaders as a frontier force guarding their new home. The somewhat troubled integration of the Sons of the Wolf into the society they are protecting is told in the story of five lovers representing both peoples, four of whom eventually marry.